title
Meristic analysis for Bonneville and Colorado river cutthroat trout in the state of Utah : annual report 2001
author
Array ( [0] => Thron, Jason E. [1] => Miller, Peggy A. )
abstract
UDWR Publication Number 02-15
date
2002-01-01
organization
Utah. Division of Wildlife Resources
species
Array ( [0] => Not Specified )
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https://grey-lit.s3.wasabisys.com/meristic-analysis-for-bonneville-and-colorado-river-cutthroat-trout-in-the-state-of-utah-annual-repo.pdf
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N 462f1.C8.7: Mer/t)(lI MERISTIC ANALYSIS FOR BONNEVILLE AND COLORADO RIVER CUTTHROAT TROUT IN THE STATE OF UTAH ANNUAL REPORT 2001 Publication Number 02-15 Utah Department of Natural Resources Division of Wildlife Resources 1594 W. North Temple Salt Lake City, Utah Kevin K. Conway, Director MERISTIC ANALYSIS FOR BONNEVILLE AND COLORADO RIVER CUTTHROAT TROUT IN THE STATE OF UTAH ANNUAL REPORT 2001 Final Report May 2002 Prepared by: Jason E. Thron Peggy A. Miller Utah Department of Natural Resources Division of Wildlife Resources 1594 W. North Temple Salt Lake City, Utah An Equal Opportunity Employer Kevin K. Conway, Director Publication Number 02-15 1.1j , STATE OF UTAH ~ NATURAL RESOURCES .,J Division of Wildlife Resources This project was partially funded under the U.S. Fish & Wildlife Service, Sport Fish Restoration Program (Project F -44-R). Utah Division of Wildlife Resources receives Federal financ ial assistance for the U.S. Fish and Wildlife Service. Under Title VI of the Civil Rights Act of 1964, Section 504 of the Rehabilitation Act of 1975, Title IX of the Education Amendments of 1972, the U.S. Department of the Interior and its bureaus prohibit discrimination on the basis of race, color, national origin, age, disability or sex (in education programs). If you believe that you have been discriminated against in any program, activity or faci lity, or if you desire further information please write to: The U.S. Fish and Wildlife Service Office for Diversity and Civil Rights Programs-External Programs 4040 North Fairfax Drive, Suite 130 Arlingron, V A 22203 11 TABLE OF CONTENTS INTRODUCTION ....... .......... ............ .. ........ ................ ........... ... .... ..... ... ........ ... ... .. ... ........ .. . 1 METHODS .......................................................................................................................... 1 RESULTS ............................................................................................................................ 2 Putative Bonneville Cutthroat Trout .. ................................ ..................................... 5 Putative Colorado River Cutthroat Trout ........ ................. .. ............ ............. .... ..... 10 DISCUSSION .................................................................................................................... 14 LITERATURE CITED ........ ................................. ..... ... .......................................... .... ....... 16 APPENDICES ................................................................................................................... 17 Appendix A. Character Ranges ............................................................................ 18 Appendix B. Putative Bonneville Cutthroat Trout Data .................... .. ................ 19 Appendix C. Putative Colorado River Cutthroat Trout Data .............................. .39 LIST OF TABLES Table 1. Summary data for putative Bonneville cutthroat trout populations ...................... 3 Table 2. Summary data for putative Colorado River cutthroat trout populations ............. .4 III Introduction Allendorf et al. (2001) describe hybridization as the "inbreeding of individuals from genetically distinct populations, regardless of the taxonomic status of the populations" (p. 621). For hybridization to occur species must exhibit some degree of spatial and temporal overlap during the reproductive portion of their life history. Geological and environmental barriers may prevent spatial overlap and therefore hybridization. Natural causes or human influence can stimulate hybridization. Removal of a geological barrier, a change in an environmental condition or stocking of nonnatives may allow for species to combine and potentially hybridize. Introgressive hybridization with introduced nonnative salmonids has been identified as a concern for the native cutthroat populations in Utah. Utah has three subspecies of cutthroat trout, the Bonneville cutthroat trout (Oncorhynchus clarki utah), the Colorado River cutthroat trout (Oncorhynchus clarki pleuriticus), and the Yellowstone cutthroat trout (Oncorhynchus clarki bouvieri). In Utah, Colorado River cutthroat trout (CRCT) were historically found in the tributaries of the Colorado River and the Green River. Bonneville cutthroat trout (BCT) were found throughout the Bonneville Basin. Yellowstone cutthroat trout were found in the Raft River drainage in the northwest corner of Utah. Nonnative salmonids stocked in Utah have had a negative effect on native cutthroat trout. The Proposed Policy on the Treatment of lntercrosses and Intercross Progeny (50 CFR Part 424, 61 FR 26) states that conserving species under the Endangered Species Act extends to hybrids, if the progeny share traits that characterize the taxon of the listed parent, and if the progeny more closely resembles the listed parents taxon than an entity intermediate between it and the other known or suspected nonlisted parent stock (Toline and Lentsch 1999). This policy has not been finalized, but U.S. Fish and Wildlife Service supports the draft. Meristic analysis is a tool that is used to identify individuals within a population that exhibit physical characteristics resembling cutthroat trout or an intermediate between cutthroat trout and rainbow trout (Oncorhynchus mykiss). The purpose for conducting meristic analysis on populations of cutthroat trout is to assist in determining the extent of hybridization in a given popUlation. The extent of hybridization in cutthroat trout populations is being further evaluated through a review of historic stocking records and by molecular anal ysis. Methods Current methods used are described by Hudson and Davis (2000). Scales above the lateral line, lateral line scales, basi branchial teeth, and pyloric caeca character counts were used in a hybrid index formula developed by Hubbs and Kuronuma (1942) and modified by Gilbert (1978) to determine extent of hybridization. lndices for each character, an average index of the character indices for each fish, and a population average index were determined. The hybrid index formula is as follows: where, I. is the index value for fish., U. is the character count for fish., MI is the adjusted mean of the character range for species one, and M2 is the mean of the character range for species two. I. will determine if a character from fish. more closely resembles species one (I. between o and 30), species two (I. between 70 and 100), or a hybridization of one and two (I. between 30 and 70) (Mayhew, 1983). Zero represents purity in species one and 100 represents purity in species two. I. values less than zero or greater than one hundred occur when character values fall beyond the range of the adjusted mean values for species one and two. The character counts beyond the range of the adjusted mean values for species one and two have the same impact as the adjusted mean values, so the negative Ia values are converted to zero and those over one hundred are convelted to one hundred. Adjusted mean values were used to calculate index values in the 2001 report to maintain consistency with the 2000 report. The character range as described in Trotter (1987) and Behnke (1992), mean value of the character range, and adjusted mean value of the character range for BCT, CRCT and rainbow trout are listed in Appendix A. The mean value for a character was adjusted to ensure that the character range for species one found in the literature fell within the 0 to 30 index range. For example if a BCT had a lateral line scale count of 140 (Ua ) , which is the low value within the character range for BCT, and the mean values found in the literature were used in the index formula for the BCT character range (MI=160) and for rainbow trout character range (M2~130), the index value would be 66 indicating a hybrid range character rather than a character within the BCT range. Using the adjusted mean for lateral line scales (MI= 144.2) rather than the mean (MI=160) in the index formula yields an index of 30, which is within the range for BCT. Results A summary of the index values for putative BCT and putative CRCT are found in Tables 1 and 2, respectively. One population average index out of 19 putative BCT populations was within the hybrid range. Eighteen population average indices were within the range for BCT. One population average index out of 18 putative CRCT populations was also within the hybrid range. Seventeen population average indices were within the range for CRCT. The number and proportion of average indices for individual fish in a population within the hybrid and rainbow trout range were reported. Out of 19 BCT trout populations, five populations had all individual fish average indices within the range for BCT. Seven populations had 10 percent of the individual fish average indices within the hybrid or 2 Table I . Summary data for putative Bonneville cutthroat trout populations. Water ID Stream name _ 11_~ AR~AN 4 _ ~An~_~_4_~_ I ,O IV AP 030D OI A Wolf Creek South Fork IV AI' 030D 02 IV AI' 130 IV AI' 140 IV AP 140 IV AI' 230 01 IV AP 230B 01 IV AQ200A Not Available Not A vai lab fe V AF 040A 01 V AK 020 U 0 1 V AK 020G 01 V AK 04011 01 VI AA 5 10GO I VI AS 050A 2 VI AU 070A VI AS 070A VI All 070A Where, Ogden River North Fork I-ligh Line Creek Deep Creek North Fork Deep Creek South Fork Chalk Creek Mcd-Low i-lufTCrcek Birch Creck High Pctectncct Creek Right Fork Wimmer Ran ch Creek Little Deer Creek South Fork Little Diamond Creek Cottonwood Creek Clear Creek Righ i Fork Deep Creek I3irch Creek North Creek North Fork Lower North Creek North Fork Middle North Creek North Fork Upper 0 = Bonneville cUllhroal trout ,30 - 70 Ilybrid '1 00 :.!~HinbO\y"!rol~ 1.9 24.9 28.5 7.9 3~,iI 0.0 8.3 13. 1 27 .7 8.3 2.7 16.2 34 .1 4.3 1.9 15.4 26.4 8.9 29.3 1.9 3.5 0.0 0.0 31.7 0.0 6.7 8.3 32.5 0.0 15.0 28. 1 11.0 40.7 13.2 13.2 7.9 12.1 9.9 4.3 6.7 0.0 x x 39.6 27.7 11.5 11 .5 x x 4.8 5.9 0.0 24.2 5. 1 15.8 15.4 10.7 17.4 28.1 26.1 10.2 5.9 7.5 42 .8 3.5 19.8 5.9 25.3 1.1 19.4 5.9 25.8 8.3 0.0 25.3 12 .3 30.9 10.7 5.9 333 9.9 99 15.0 22.2 12.3 5.9 12.1 41 2 11.5 1.9 33.3 16.6 )5 .7 13.9 7.5 11. 5 38.9 0.0 4.3 8.3 2 1.4 6.7 1.1 6.7 6.7 .15 .7 6.7 115 15.8 14 .6 13.1 12.3 11.5 13.1 18.6 .".- - 583 58.4 782 ;:-;, li,.2 30. 1 56.1 34.9 1.1 1.9 1.9 39 (, 26.6 11.5 4.3 6.7 7.5 3.5 2.7 Ave Ia = The average character index for the individual fi sh within the sample #1-11 = The nllmber of hybrid and rainbow troul range individual average indices %1-11 = percent of hybrid and rainbow trout range individual fish average indices #HC =The number of hybrid and rainbow trout range individual character indices for the sample #TC = The total number of individual characters for the sample %HC = The percent of hybrid and rainbow Irout range individual character indices for the sample x = Ind ividua ls too small , damaged, or absent for accurate results 3 4.3 13.4 22.9 x x 28.5 9. 1 3.5 33.3 3.5 31.7 67.0 26.2 11.5 15.0 27.0 31.4 30.9 5.9 0.0 5.1 4.3 24.5 15.0 19.7 x x x x 34.1 9.5 39.6 19.0 19.8 9.9 16.2 3.5 2 1.8 7.9 3.5 8.3 51.7 10.7 7.5 19.4 16.6 10.7 38.9 9. 1 15.41 5.4 55 .0 49.'1 11.8 9. 1 22.6 3.5 mean #1-11 4.8 15.8 18.4 5.7 25.1 12.8 18.4 13 .2 16.7 12.6 14.9 25.2 2 1.1 8.5 14.8 15.6 5 1.8 16.7 9.7 o I o 2 2 4 3 o 2 o 9 3 o %HI ! HC/!TC %HC 10.0 0.0 10.0 0.0 40.0 10.0 10.0 10.0 10.0 10.0 20.0 40.0 30.0 0.0 20.0 0.0 90 .0 30.0 0.0 1/40 8/40 10127 1117 7119 6/40 15/40 6140 9/38 7/38 10/40 13/40 13/40 7140 8/40 11/40 24/39 9/39 3140 2.5 20.0 37.0 5.9 36.8 15.0 37.5 15.0 23.7 18.4 25.0 32.5 32.5 17.5 20.0 27.5 6 1.5 23. 1 7.5 Table 2. SUlllmary data for putative Colorado River cutthroat trout populations. Water I)) Stream name _~_~_~_~_~_4An~AR~An~_ I ,. I AJ 110 C Boulder Creek East Fork I I30 03 Indian Creek I I3Q 050 0 I Roc Creek Station A I I3Q 050A 0 I Taylor Creek I DQ 070 03 l.a Sal Creek I BQ Not Assigned La Sal Creek Ditch Main Divers ion II AI 130M 05 Lake Canyon Creek I[ AI 130N Nuck Woodward Creek II AI 139M 0 1 01 Scad Valley Creck II AK 100A 01 Gordon Creek North Fork II AK 170A OI KyullcRcservoir Crcck II i\K 190 13 01 AOI White River Middle Fork II DB Not Assigned Steer Gulch JIIJE 02013 04 01 Garfield Creek II DE 0200 01 Brown Ouck Creek II BE 1501 01 Duchesne River Little west Fork IIBEI50FOI Val Creek II CJ 040 13 Beaver Creek Middle Fork Where, 0= Colomdo River Clll1hroat trout 30 - 70 ~ Ilybrid 00 .. Raillbow \rOU\ •. --,w. 0.0 1.1 14.7 38.7 4.5 IQ7 5.5 5~ 3~ 3.4 53 4.7 4.1 12.5 27.5 20.2 0.0 0.0 12.0 14.4 8.9 11~.4 6.7 4.7 1.1 0.0 24.3 18.4 18.8 5.6 0.0 0.0 0.0 0.3 4.1 3.6 3.6 18.3 1.1 1.1 0.0 42.2 3.5 23.5 4.2 15.9 7.6 17.8 2.0 23.1 11.1 0.0 0.0 2.4 3.6 0.0 14 .1 115.6 5.1 5.9 7.5 2.9 1.8 0.9 14.1 9.9 13.6 43 17 .5 ) 1.9 28.6 19.1 13.3 15.3 67.9 .. 19,2 25.0 11.6 9.1 11.5 25.1 12.0 5.3 13.5 3.3 0.0 0.0 '12.7 0.7 0.0 Ave la = The average character index for the individual fi sh within the sample #1-11 = The number of hybrid and rainbow trout range individual average indices %H I = percent of hybrid and rainbow trout range individual fi sh average indices 0.0 0.0 1.5 13.8 20 .4 17.2 4.3 30.9 8.3 0.5 0.0 0.3 0.0 7.0 20.2 11.7 11.2 ~4 .9 20.0 12 .2 3.5 40.8 19.7 22.0 23.2 29.4 9.3 30.2 20.9 3. 1 45.2 61.7 .41.7 5.1 8.3 15 .8 6.8 12.2 5.9 20.0 25.2 16.5 11.8 16.5 17.4 x x x x x x 25.9 x x #HC =The number of hybrid and rainbow trout range individual character indices for the sample HTC = The total number of individual characters for the sample %HC = The percent of hybrid and rainbow trout range individual character indices for the sample x = Individuals too small , damaged, or absent for accurate results 4 1.8 14.4 3.5 34.8 0.0 7.5 4.5 24.4 5.5 323 9.9 5.4 2 1.1 4.2 9.6 x x x 0.3 27.2 1.1 1.1 7.1 33.9 11.9 24.5 7.6 18.0 10.5 3.7 5.3 7.8 12.4 x x x mean II I-II %111 HIIC/IITe %HC 1.2 22.4 7.5 5.9 4.4 18.5 9.0 27.2 12.7 17.5 40.9 10.4 7.5 17.7 12.2 0.7 8.6 6.1 o 2 1 o 3 o 2 o 2 6 o o o o o o 0.0 20.0 10.0 10.0 0.0 30.0 0.0 20.0 0.0 20.0 60.0 0.0 0.0 0.0 0.0 0.0 20.0 0.0 0/39 12/40 3/40 1/40 1/40 7/40 5/40 14/36 5138 10/40 16/37 5140 2/40 9/40 9/40 0/20 1/14 1/23 0.0 30.0 7.5 2.5 2.5 17.5 12.5 38.9 13.2 25.0 43.2 12.5 5.0 22.5 22.5 0.0 7.1 4.3 rainbow trout range, two populations had 20 percent, two had 30 percent, two had 40 percent and one had 90 percent of the individual fish average indices in the population within the hybrid and rainbow trout range. Two BCT populations had small sample sizes and those data should be interpreted with care. Out of 18 CRCT populations, 10 populations had all individual fish average indices within the range for the CRCT. Two populations had 10 percent of the individual fi sh average indices within the hybrid and rainbow trout range, four populations had 20 percent, one had 30 percent, and one population had 60 percent within the hybrid and rainbow trout range. Three CRCT populations had small sample sizes. The number and proportion of hybrid and rainbow trout range character indices in a population were also reported. The proportion of hybrid and rainbow trout range character indices is the total number of hybrid and rainbow trout range character indices in a population divided by the total number of characters counted in a population. The total number of characters counted in a population varied between populations. The number of individual fish in a sample was not always the same and sometimes characters were too damaged to count. The proportion allows for comparison across populations, but data from populations with small sample sizes and low character counts should be interpreted with care. The results for each population are summarized below. The information has been separated into putative BCT and putative CRCT for convenience. This summary is based on the data found in Appendices B and C. Putative Bonneville Cutthroat Trout WolfCreek South Fork IV AP 030D OIA The population average index (4.8) for WolfCreek -South Fork was within the range for BCT. The individual fish average indices and character indices for nine out of 10 fish were within the range for BCT. One fish had an individual fish average index (31.7) within the hybrid range and a character index for basi branchial teeth within the range for rainbow trout. Of the characters counted, 2.5% of the character indices were within the hybrid range. Odgen River - North Fork (high) IV AP 030D 02 The population index average (15.8) for Ogden River - North Fork (high) was within the range for BCT. The individual fish average indices for nine out of 10 fish were within the range for BeT. One fish had an individual fish average index (32.5) within the hybrid range and a rainbow trout range character index for basi branchial teeth. Six additional fish had at least one hybrid or rainbow trout range character index. Of the 5 characters counted, 20% of the character indices were within the hybrid or rainbow trout range. Line Creek IV AP 130 The population index average (18.4) for Line Creek was within the range for BCT. The individual fish average indices for nine out of 10 fish were also within the range for BCT. One fish had an individual fish average index (40.7) and two character indices (scales above the lateral line and basibranchial teeth) within the hybrid range. Seven addition fish had at least one character index within the hybrid range. The fish did not preserve well, so counts were not obtained for basbranchial teeth in three fi sh and pyloric caeca in all 10 fish. Of the characters counted, 37% of the character indices were within the hybrid or rainbow trout range. Deep Creek North Fork IV AP 140 Five fish were analyzed for Deep Creek - North Fork. The population average index (5.7) and the individual fish average indices were all within the range for BCT. Scales above the lateral line and the lateral line scales were not counted in one fish due to size. Pyloric caeca was not counted in another fish. Only one individual character index for scales above the lateral line fell into the hybrid range. All other countable character indices were in the range for BCT. Of the characters counted, 5.9% of the character indices were within the hybrid range. Deep Creek South Fork IV AP 140 Five fi sh were analyzed fo r Deep Creek South Fork. The population average index (25 .1 ) was within the range for BCT. Two out of five fi sh had hybrid range individual fi sh average indices (35.4 and 39.6). Three addition fi sh had one character range index within the hybrid or rainbow trout range. Of the characters cOll.'1ted, 36.8% of the character indices were within the hybrid or rainbow trout range. Chalk Creek (medium-low) IV AP230 01 The population average index (12.8) for Chalk Creek (medium-low) was within the range for BCT. Nine out of 10 fish had individual fish average indices within the BCT 6 range. One fish had a hybrid range individual fish average index (34.1), a rainbow trout range character index for pyloric caeca and a hybrid range character index for basi branchial teeth. Three additional fish had at least one character index in the hybrid or rainbow trout range. Of the characters counted, 15% of the character indices were within the hybrid or rainbow trout range. Huff Creek IV AP 230B 01 The population average index (18.4) for Huff Creek was within the range for BCT. The individual fish average indices for nine out of 10 fish were also within the BCT range. One fish had a hybrid range individual fish average index (39.6). All 10 fish had at least one hybrid or rainbow trout range character index. Nine out of 10 fish had hybrid range character indices for scales above the lateral line. Of the characters counted, 37.5% of the character indices were within the hybrid or rainbow trout range. Birch Creek High IVAQ200A The population average index (13.2) for Birch Creek High was within the range for BCT range. The individual tish average indices for nine out of 10 fish were also within the range for BCT. The fish with the individual fish average index (42.8) within the hybrid range had a hybrid range character for basi branchial teeth and a rainbow trout range character for pyloric caeca. Five out of 10 fish were within the BCT range for all character indices. Four other fish had character indices within the hybrid or rainbow trout range. Of the characters counted, 15% of the character indices were within the hybrid or rainbow trout range. Peteetneet Creek RighI Fork Spanish Fork Drainage The population average index (16.7) for Peteetneet Creek Right Fork was within the range for BCT. The individual fish average indices for nine out of 10 fish were also within the range for BCT. The fish with the individual fish average index (33.3) within the hybrid range had a hybrid range character index for scales above the lateral line and a rainbow trout range character index for basibranchial teeth. Six other fish also had hybrid or rainbow trout range character indices. Of the characters counted, 23.7% of the character indices were within the hybrid or rainbow trout range. 7 Wimmer Ranch Creek None The population average index (12.6) for Wimmer Ranch Creek was within the range for BCT. The individual fish average indices for nine out of 10 fish were also within the BCT range. The fish with the individual fish average index (30.9) within the hybrid range had a rainbow trout range character index for basibranchial teeth. Five other fish had character indices within the hybrid range. Of the characters counted, 18.4% of the character indices were within the hybrid range. Little Deer Creek South Fork V AF 040A 01 The population average index (14.9) for Little Deer Creek South Fork was within the range for BCT. The individual fish average indices for eight out of 10 fish were also within the BCT range. Of the two fish that were within the hybrid range for individual fish average indices, one (33.3) had a hybrid range character index for scales above the lateral line and a rainbow trout range character index for basi branchial teeth. The other (31. 7) had a rainbow trout range character index for basibranchial teeth. Six additional fish had at least one character index within the hybrid range. Of the characters counted, 25% of the character indices were within the hybrid or rainbow trout range. Little Diamond Creek V AK 020B 01 The population average index (25.2) for Little Diamond Creek was within the range for BCT. The individual fish average indices for six out of 10 were also within the BCT range. Four fish had hybrid range individual fish average indices (33.3, 41.2, 51.7, 67.0). Eight fish had at least one character index within the hybrid or rainbow troUl range. Of the characters counted, 32.5% of the character indices were within the hybrid or rainbow trout range. Cottonwood Creek V AK020G 01 The population average index (21.1) for Cottonwood Creek was within the range for BCT. The individual fish average indices for seven out of 10 fish were also within the BCT range. Three fish had individual fish average indices within the hybrid range (34.1 , 35.7, and 38.9). All three fish had hybrid range character indices for scales above the lateral line and rainbow trout range character indices for basi branchial teeth. Five additional fish had at least one character index within the hybrid or rainbow trout range. Of the characters counted, 32.5% of the character indices were within the hybrid or rainbow trout range. 8 Clear Creek Right Fork V AK040HOI The population average index (8.5) for Clear Creek Right Fork was within the range for BCT. All 10 fish were also within the BCT range for individual fish average indices. All character indices for five fish were within the range for BCT. The other five fish had at least one hybrid range character index. Of the characters counted, 17.5% of the character indices were within the hybrid or rainbow trout range. Deep Creek VI AA 510G The population average index (14.8) for Deep Creek was within the range for BCT. Eight out of 10 fish were also in the BCT range for individual fish average indices. Two fish had hybrid range individual fish average indices (38.9, 35.7). The character indices for scales above the lateral line were within the hybrid range and the character indices for basi branchial teeth were within the rainbow trout range for both fish. Four additional fish had one hybrid range character index each. Of the characters counted, 20% of the character indices were within the hybrid range. Birch Creek VI AB 050A 2 The population average index (15.6) for Birch Creek was within the range for BCT. All 10 fish were also within the BCT range for individual fish average indices. All 10 fish had at least one character index within the hybrid or rainbow trout range. Of the characters counted, 27.5% of the character indices were within the hybrid or rainbow trout range. North Creek Lower North Fork VIAB 070 A The population average index (51.8) for North Creek Lower North Fork was within the hybrid range. One out of 10 fish had an individual fish average index (26.4) within the BCT range, seven had individual fish average indices (58.3, 58.4, 30.1, 56.1, 31.4, 55.0, 49.4) within the hybrid range and two were within the rainbow trout range (78 .2, 74.2). All fish had at least one hybrid or rainbow trout range character index. Of the characters counted, 61.5% of the character indices were within the hybrid or rainbow trout range. 9 North Creek North Fork Middle Station VIAB 070 A The population average index (16.7) for North Creek North Fork Middle Station was within the range for BCT. The individual fish average indices for seven out of 10 fish were also within the BCT range. Three fish were in the hybrid range for individual fi sh average indices (34.9, 39.6, and 30.9). Four fish had all BCT range character indices. The additional six fish had at least one character index within the hybrid or rainbow trout range. Of the characters counted, 23.1 % of the character indices were within the hybrid or rainbow trout range. North Creek North Fork Upper Station VIAB 070A The population average index (9.7) for North Creek North Fork Upper Station was within the range for BCT. All 10 fish were also within the BCT range for individual fish average indices. Seven fi sh had all BCT range character indices. Three fish had one character index within the hybrid range or the rainbow trout range. Of the characters counted, 7.5% of the character indices were within the hybrid or rainbow trout range. Putative Colorado River Cutthroat Trout Boulder Creek East Fork I AJ IIOC The population average index (1.2) for Boulder Creek East Fork was within the range for CRCT. All the individual fi sh average indices and all the character indices for each fi sh were also within the range for CRCT. Pyloric caeca were not counted for one fish. Indian Creek I BG03 The population average index (22.4) for Indian Creek was within the range for CRCT. Eight out of 10 fish were within the CRCT range for individual fish average indices. Two fi sh were within the hybrid range for individual fish average indices (38.7, 45.6). Nine fish had at least one hybrid or rainbow trout range character index. Of the characters counted, 30% of the character indices were within the hybrid or rainbow trout range . 10 Roc Creek I BQ 050 01 The population average index (7.S) for Roc Creek was within the range for CRCT. The individual fish average indices for nine out of 10 fish were also within the CRCT. One fish had a hybrid range individual fish average index (30.9) with a rainbow trout range character index for basibranchial teeth. Two fish had hybrid range character indices for scales above the lateral line. Of the characters counted, 7.S% of the character indices were within the hybrid or rainbow trout range. Taylor Creek I BQ 050AOI The population average index (S.9) for Taylor Creek was within the range for CRCT. Nine out of 10 fish were within the CRCT range for individual fish average indices and individual character indices. One fish had an individual fish average index (34.8) within the hybrid range and a rainbow trout range individual character index for basi branchial teeth. Of the characters counted, 2.S% of the character indices were within the rainbow trout range. La Sal Creek (section 3) I BQ 070 03 The population average index (4.4) for La Sal Creek (section 3) was within the range for CRCT. All 10 fish were also within the CRCT range for individual fish average indices. Only one fish had a hybrid range character index for basi branchial teeth. All other character indices were within the CRCT range. Of the characters counted, 2.S% of the character indices were within the hybrid range. La Sal Creek Ditch - Main Diversion Water ID not assigned to this stream The La Sal Creek Ditch - Main Diversion population average index (18.S) was within the range for CRCT. The individual fish average indices for seven out of 10 fish were also within the CRCT range. Three fish had hybrid range individual fish average indices (42.2,44.9, and 33.9) with hybrid range character indices for scales above the lateral line and rainbow troUl range character indices for basi branchial teeth. One other fi sh had a hybrid range character index for scales above the lateral line. Of the characters counted, 17.S% of the character indices were within the hybrid or rainbow trout range. II Lake Canyon Creek II AI 130M 05 The Lake Canyon Creek population average index (9.0) was within the range for CRCT. All 10 fish were also within the CRCT range for individual fish average indices. Four fish had one hybrid range character index. Of the characters counted, 12.5% of the character indices were within the hybrid range. Nuck Woodward Creek II AI l30N The Nuck Woodward population average index (27.2) was within the range for CRCT. Eight out of 10 fish were within the CRCT range for individual fish average indices. Two fish had individual fish average indices (51.9, 40.8) within the hybrid range. Nine fish had at least one hybrid or rainbow trout range character index. Basibranchial teeth were not counted in four fish. Of the characters counted, 38.9% of the character indices were within the hybrid or rainbow trout range. Scad Valley Creek II AI 139M 01 01 The Scad Valley Creek population average index (12.7) was within the range for CRCT. All 10 fish were also within the CRCT range for individual fi sh average indices. Four fi sh had at least one hybrid or rainbow troUl range character index. Basibranchial teeth were not counted in two fish due to damage. Of the characters counted, 13.2% of the character indices were within the hybrid or rainbow trout range. Gordon Creek - North Fork II AK 100A 01 The Gordon Creek - North Fork population average index (17.5) was within the range for CRCT. Eight out of 10 fish were also within the CRCT range for individual fish average indices. Two fish were in the hybrid range (30.2, 32.3) for individual fi sh average indices. Nine fish had at least one hybrid or rainbow troUl range character index. Of the characters counted, 25% of the character indices were within the hybrid or rainbow trout range. Kyune Reservoir Creek II AK 170A 01 The Kyune Reservoir Creek population average index (40.9) was within the hybrid range. Four out of 10 fi sh for individual fi sh average indices and all individual character 12 indices were within the CRCT range. Four fish had individual fish average indices (67.9, 45.2, 61.7, and 41.7) within the hybrid range and two fish had individual fish average indices (76.4, 79.0) within the rainbow trout range. Five fish had at least one hybrid or rainbow trout character range index. Pyloric caeca were not counted in three fish. Of the characters counted, 43.2% of the character indices were within the hybrid or rainbow trout range. White River - Middle Fork • II AK 190 BOI AOI '(This population listed in the 2000 report (Hudson and Davis 2000), was actually White River Right Fork Upper II AK 190A) The White River - Middle Fork population average index (10.4) was within the range for CRCT. All 10 individual fish average indices were also within the CRCT range. Five fish had at least one character index within the hybrid or rainbow trout range. Of the characters counted, 12.5% of the character indices were within the hybrid or rainbow trout range. Steer Gulch (West Willow Creek) None Available The Steer Gulch (West Willow Creek) population average index (7.5) was within the range for CRCT. All 10 individual fish average indices were also within the CRCT range. Only one fish had hybrid range character indices for scales above the lateral line and lateral line scales. Of the characters counted, 5% of the character indices were within the hybrid range. Garfield Creek II BE 020B 04 01 The Garfield Creek population average index (17.7) was within the range for CRCT. All 10 individual fish average indices were also within the CRCT range. Seven fish had at least one hybrid or rainbow trout range character index. Of the characters counted, 22.5% of the character indices were within the hybrid or rainbow trout range. Brown Duck Creek II BE 020D 01 The Brown Duck Creek population average index (12.2) was within the range for CRCT. All 10 individual fish average indices were also within the CRCT range. Two fish had all CRCT range character indices. Eight fish had at least one hybrid range character 13 index. Of the characters counted, 22.5% of the character indices were within the hybrid range. Little West Fork - Duchesne River II BE 1501 01 The Little West Fork - Duchesne River population average index (0.7) was within the range for CRCT. Only five fish were available for meristic analysis. All five individual fish average indices and individual character indices were within the CRCT range. Vat Creek II BE 150FO I Five fish were available for meristic analysis in the Vat Creek sample. Pyloric caeca were not counted in one fish, pyloric caeca and basi branchial teeth were not counted in a second fish, and lateral line scales, pyloric caeca and basi branchial teeth were not counted in a third fish. Due to the low number of fish within the sanlple and the low number of characters counted per fish, care should be taken in interpreting these data. The population average index (8.6) was within the range for CRCT. The individual fish average indices for four fish were within the CRCT character range. The individual fi sh average index (42.7) for one fish was within the hybrid range, but the individual fish average index was based on one character index, the scales above the lateral line. Of the characters counted, 7.1 % ofthe character indices were within the hybrid range. Beaver Creek - Middle Fork II CJ 040B The Beaver Creek - Middle Fork population average index (6.1) was within the range for CRCT. Six fish were available for meristic analysis. All six individual fish average indices were also within the CRCT range. One fish had a rainbow trout character index for basi branchial teeth. All other character indices were within the CRCT range. Of the characters counted, 4.3 % of the character indices were wi thin the rainbow trout range. Discussion As noted in the 2000 Annual Report (Hudson and Davis 2000), there are limitations associated with interpreting the results using the hybrid index formula. The hybrid index does not take into account hybridization between cutthroat subspecies and may bias the rainbow and cutthroat trout hybridization results. Some character count ranges overlap between rainbow and cutthroat trout and between cutthroat trout subspecies. 14 An individual with an index close to zero is no more pure than an individual with an index close to 30. The index is an indication of how close the character counts are to the adjusted mean of the character counts found by Trotter (1987) and Behnke (1992). Individuals with hybrid range average indices may have several hybrid range character indices or one rainbow trout range character index and several cutthroat rrout range character indices close to 30. With the limitations taken into consideration, the hybrid index results indicate 17 of the populations analyzed in 200 I have indices within the range for BCT and 18 of the populations analyzed in 2001 have indices within the range for CRCT. If genetics results verify these results, those populations will be available for the conservation of BCT and CRCT in Utah. 15 Literature Cited Allendorf, F. W., R. F. Leary, P. Spruell, and 1. K. Wenburg. 2001. The problems with hybrids: setting conservation guidelines. Trends in Ecology & Evolution. Vol 16 No. 11 November 200 1: 613-622. Behnke, R. J. 1992. Native trout of western North America. American Fisheries Society. Monograph No.6 Bethesda, MD. Gilbert, C. R. 1978. The nominal North American cyprinid fish Notropis henryi interpreted as an intergeneric hybrid, Clinostomus fllnduloides X Nocomis leptocephalus. Copeia 1978: 177-181. Hubbs, C. 1. and K. Kuronwna. 1942. Hybridization in nature between two genera of flounders in Japan. Ibid. 27:267-306. Hudson, J. M. and C. J. Davis. 2000. Meristic analysis results for Bonneville and BCT in the state of Utah, Annual report November 2000. Utah Department of Natural Resources, Division of Wildlife Resources. Publication Number 00-34. Pp.70. Mayhew, D. A. 1983. A new hybrid cross, Notropis atherinoides X Notropis volucelills (P isces: Cyprinidae), from the lower Monongahela River, western Pennsylvania. Copeia 1983:1077-1082. Toline, C. A. and 1. D. Lentsch. 1999. Guidelines and protocols for identification and designation of populations of native cutthroat trout. Final report to the Utah Division of Wildlife Resources. Pp. 41 . Trotter, P. C. 1987. Cutthroat: native trout of the west. Colorado Associated University Press. Boulder, CO. 16 Appendices 17 Appendix A. Character ranges fo r Bonneville cutthroat trout, Colorado River cutthroat trout, and rainbow trout. Bonneville cutthroat trout Colorado River cutthroat trout Rainbow trout Range Adjusted Range Adjusted Range Adjusted Character (J i terature) Mean Mean (literature) Mean Mean (I i terature) Mean Mean Scales above the lateral line 38-46 42 42.7 38-48 43 42.7 26-28 27 27 Lateral line scales 140-180 160 144.2 170-205 187 187 120-140 130 130 Pyloric caeca 25-55 40 48 24-45 35 41 50-60 55 55 Basibranchial teeth up to 90 6 3 yes 6 3 no 0 0 ---- 18 Appendix B 19 Appendix 8. Putative Bonneville cutthroat trout Population: Water 10: Wolf Creek - South Fork IV AP 0300 OIA Population Average = 4.8 Character R B U, Scales Above Lat. Line 27.0 42.7 41.5 Lateral Line Scales 130.0 144.2 163.5 G ill Rakers 20.5 18.5 19.0 Basibranchial Tecth 0.0 3.0 3.0 Pyloric Caeca 55.0 48.0 40.0 Ave I, = Character R B U, Sca les Above Lat. Line 27.0 42.7 38.5 Lateral Line Scales 130.0 144.2 156.5 Gill Rakers 20.5 18.5 17.0 Basibranchial Teeth 0.0 3.0 0.0 Pyloric Caeca 55.0 48.0 37.0 Ave 1,= I, = 100 *[(U,-B)/(R-B)] where, R = th e mean or tile character range for rainbow trout I, U, 7.6 41.5 0.0 189.0 x 18.0 0.0 5.0 0.0 30.0 1.9 Ave I, = I, U, 26.8 45.0 0.0 170.0 , 17.0 [,!00,2. 6.0 0.0 43.0 31.7 Ave 17 = B = the adjusted mean of the character range for Bonneville cutthroat trout Ua = the raw character count for each fish I, = the character hybrid index for each fish Ave 18 = the average or the character hybrid indices for each fish 0 = Bonneville 30 . 70 a Ilybrid '120 Ra~ 20 I, U, I, U, I, U, I, 7.6 40.5 14.0 42.5 0.0 42.5 0.0 0.0 179.5 0.0 187.5 0.0 161.0 0.0 , x 19.0 x 17.0 x 18.0 x 0.0 3.0 0.0 3.0 0.0 4.0 0.0 0.0 35.0 0.0 36.0 0.0 38.0 0.0 1.9 Ave I, = 3.5 Ave I, = 0.0 Ave 1,= 0.0 I, U, I, U9 I, U,o 1'0 0.0 40.0 17.2 43.5 0.0 39.5 2004 0.0 175.0 0.0 186.5 0.0 178.5 0.0 x 19.0 x 20.0 x 18.0 x 0.0 4.0 0.0 5.0 0.0 7.0 0.0 0.0 39.0 0.0 38.0 0.0 36.0 0.0 0.0 Ave 18 = 4.3 Ave I, = 0.0 Avc11O = 5.1 Appendix B. Putative Bonneville cutthroat trout Population: Water lD: Population Average = Characler Ogden River - North Fot'k (high) IV AP 030D 02 15.8 R B U, I, U, I, --------- -- ----- -- - - - - - - -- - Scales Above Lal. Line 27.0 42.7 37.5 33.1 38.5 Laleral Line Scales 130.0 144.2 15 1.5 0.0 176,5 Gi ll Rakers 20.5 18.5 18.0 x 21.0 Basibranchial Teelh 0.0 3.0 1.0 66.7 4,0 Pyloric Caeca 55.0 48,0 40.0 0.0 47.0 Ave " ~ 24.9 Ave " ~ Character R B U, I, U, Scales Above Lal. Line 27.0 42.7 38.5 26,8 38.5 Lateral Line Scales 130,0 144.2 156.5 0,0 179.5 Gill Rakers 20.5 18.5 18,0 x 19.0 Basibranchial Teeth 0.0 3.0 2.0 33.3 3.0 Pyloric Caeca 55.0 48.0 43 .0 0.0 54.0 Ave 16= 15.0 Ave I , ~ I, ~ 100 *[(U.-B)/(R-B)] where, R = the mean orthe character range for rainbow trout B ~ Ihe adjusled mean of the character range for Bonneville cullhroallroul Ua = the raw character count for each fish I, ~ the characler hybrid index for each fi sh Ave I, ~ Ihe average oflhe characler hybrid indices for each fi sh o ~ Bonneville 30 - 70 /lybrid I OQ "'_Rainbow 26.8 0.0 x 0.0 0.0 6.7 I, 26.8 0,0 x 0.0 t tt? 28.1 21 U, I, U, I, U, I, ---- ---- -- --- 37.5 33.1 38.0 29.9 43.0 0.0 165.0 0.0 159.0 0.0 157.0 0.0 17.0 x 21.0 x 19.0 x 8.0 0.0 0.0 0 60.0' 8.0 0.0 44,0 0.0 38.0 0.0 42,0 0.0 Ave I, ~ 8.3 Ave I, ~ 32.5 AveJ, ~ 0.0 U, I, U, I, U,o 1'0 39.5 20.4 40,0 17.2 38.5 26.8 16/.5 0.0 169.0 0.0 173.0 0,0 2 1.0 x 17.0 x 2 1.0 x 2.0 33.3 8.0 0.0 5.0 0.0 I 45.0 0.0 36.0 0.0 53.0 flM Ave 18 = 13.4 Ave I, ~ 4.3 Ave 1'0 ~ 24.5: Appendix B. Putative Bonneville cutthroat trout Population: Line Creek Water lD : IV AP 130 Population Average = 18.4 Character R B U, I, U, Scales Above Lat. Line 27.0 42.7 34.5 52.2 37.5 Lateral Line Scales 130.0 144.2 174.0 0.0 169.0 Gill Rakers 20.5 18.5 19.0 x x Basibranchial Teeth 0.0 3.0 2.0 33.3 3.0 Pyloric Caeca 55.0 48.0 x x x Ave II = 28.5 Ave 12 = Character R B U6 16 U7 Scales Above Lat. Line 27.0 42.7 39.0 23.6 37.0 Lateral Line Scales 130.0 144.2 179.0 0.0 170.0 Gill Rakers 20.5 18.5 19.0 x 19.0 Basibranchial Teeth 0.0 3.0 3.0 0.0 5.0 Pyloric Caeca 55.0 48.0 x x x Ave 16 = 7.9 Ave 17 = I, = 100 *[(U,-B)/(R-B)] where, R ::::: the mean of the character range for rainbow trout B = the adjusted mean of the character range for Bonneville cutthroat trout Ua = the raw character count for each fi sh I, = the character hybrid index for each fi sh Ave I, = the average of the character hybrid indices for each fish 0 = Bonneville 30 - 70 = Hybrid I.IQQ., ~n!!!.ll.Q~y' I, U, I, U, 14 U, I, 33. 1 34.0 , 55 .4 36 .5 39.5 36.5 39.5 0.0 159.0 0.0 172.0 0.0 191.0 0.0 x 17.0 x 20.0 x 18.0 x 0.0 1.0 t~Q~".7 _ 4.0 0.0 5.0 0.0 x x x x x x x 11.0 Ave I, = 40.7 Ave 14 = 13.2 Avcl s= 13.2 17 U, I, U. I, U,o 1'0 36.3 35.5 45 .9 38 .0 29.9 36.5 39.5 0.0 171.5 0.0 172.0 0.0 162.5 0.0 x 17.0 x 18.0 x 19.0 x 0.0 x x x x x x x x x x x x x 12.1 Ave I, = 22.9 Ave 19 = 15.0 Avc11O = 19.7 22 Appendix B. Putative Bonneville cutthroat trout Population: Water 10: Deep Creek_NF IV AP 140 l>opuIatioll Average = 5.7 Character R .. B - Scales Above Lat. Line 27.0 42.7 Lateral Line Scales 130.0 144.2 Gill Rakers 20.5 18.5 Basibranch ial Teeth 0.0 3.0 Py loric Caeca 55.0 48.0 U, - , 39.0 173.5 18.0 3.0 x I, U, . 23.6 36.5 0.0 181.5 x 2 1.0 0.0 4.0 x 40.0 Ave I, = 7.9 Ave I, = I, = 100 '[(U,-B)/(R-B)] where, R = the mean of the c B = the adjusted mean of the character range for Bonneville cutthroat trout Uft = the raw character count for each fish I, = the character hybrid index for each fish Ave J, = the average of the character hybrid indices for each fi sh 0 = Bonneville 30 - iiJ = Ilybrid 100 = Rainbow I, . U, ~ I, ~ U, . . I, . U, ~ 0 39.5 40.0 17.2 38.5 26.8 x x 0.0 181.0 0.0 178.0 0.0 x x x 20.0 x 20.0 x x x 0.0 7.0 0.0 6.0 0.0 4.0 0.0 0.0 x 0.0 43.0 0.0 46.0 0.0 9.9 Ave I, = 4.3 Ave I, = 6,7 Ave 1, = 0.0 23 Appendix B. Putative Bonneville cutthroat trout Population: Water 10: Deep CrecI{-SF IV AP 140 Population Average = 25.1 Character R o Scales Above Lat. Line 27.0 42.7 Lateral Line Scales 130.0 144.2 Gill Rakers 20.5 18.5 Basibranchial Teeth 0.0 3.0 Pyloric Caeca 55.0 48.0 U, 36.5 174.0 19.0 1.0 x I, U, 39.5 33.5 0.0 154.5 x 20.0 66.7 0.0 x 35.0 Ave I, = 35.4 Ave I, = I, = 100 ·[(U,·B)/(R-O)] where, R = the mean of Ihe c B = the adjusted mean of the character range for Bonneville Clilthroattrolit Ua = the raw character count for each fi sh I, = the character hybrid index for each fi sh Ave la = the average of the character hybrid indices for each fish o ~ Bonneville 30 - 70 = lIybrid 100 u Rainbow I, U, I, U4 14 U, I, 58.6 41.0 10.8 35.5 45 .9 35.5 45.9 0.0 180.0 0.0 155.5 0.0 158.5 0.0 x 19.0 x 20.0 x x x r lOO~ 0.0 UQQ&. 5.0 0.0 3.0 0.0 0.0 38.0 0.0 37.0 0.0 44.0 0.0 39.6 Ave I, = 27.7 Ave 14 = 11.5 Ave 1, = 11.5 24 Appendix B. Putative Bonneville cutthroat trout Population: Water 10: Chalk Creek - (medium-low) IV AP 230 01 Population Average = 12.8 Character R B U, I, -- Scales Above Lat. Line 27 .0 42.7 45.0 0.0 Lateral Line Scales 130.0 144.2 161.5 0.0 Gill Rakers 20.5 18.5 19.0 x Basibranchial Teeth 0.0 3.0 8.0 0.0 Pyloric Caeca 55.0 48.0 41.0 0.0 Ave I, = 0.0 Character R B u, I, Scales Above Lat. Line 27.0 42.7 39.5 20.4 Lateral Line Scales 130.0 144.2 150.5 0.0 Gill Rakers 20.5 18.5 21.0 x Basibranch ial Teeth 0.0 3.0 3.0 0.0 Pyloric Caeca 55.0 48.0 44.0 0.0 Ave 1, = 5.1 I, = 100 '[(U,-B)/(R-B)] where, R = the mean of the character range for rainbow trout U, 40.0 167.5 19.0 3.0 47.0 Ave 12 = U, 39.5 165.0 20.0 8.0 51.0 Ave I, = B = the adjusted mean of the character range for Bonneville cutthroat trout U, = the raw character count for each fi sh I, = the character hybrid index for each fi sh Ave I, = the average of the character hybrid indices for each fi sh 0 = Bonneville 30 - 70 ~ Ilybrid 100 =" Rai\lbow • I, 17.2 2.0 x 0.0 0.0 4.8 I , 20.4 0.0 x 0.0 42.9 15.8 25 U, I J U, I, U, I, - - ------- 39.0 23.6 43.0 0.0 36.5 39.5 173.0 0.0 174.0 0.0 156.5 0.0 18.0 x 20.0 x 21.0 x 7.0 0.0 10.0 0.0 3.0 0.0 39.0 0.0 37.0 0.0 52.0 57.1 Ave I, = 5.9 Ave I, = 0.0 Ave 1, = 24.2 U. I. U, I, U,. I,. 40.5 14.0 40.0 17.2 39.0 23.6 168.5 0.0 182.5 0.0 163.0 0.0 20.0 x 18.0 x 21.0 x 7.0 0.0 2.0 33.3 4.0 0.0 56.0 [,iOO.O 54.0 [ 85.7 49.0 14.3 Ave I. = 28.5 Ave I, = 34.1 Ave 110 = 9.5 Appendix B. Putative Bonneville cutthroat trout Population: Huffer. WaterlD: IV AP 230B 01 Population Average = 18.4 Character R B U, I, U, Scales Above Lat. Line 27.0 42.7 37.5 33.1 33.0 Laleral Line Scales 130.0 144 .2 165.0 0.0 157.0 Gill Rakers 20.5 18.5 18.0 x 17.0 Basibranchial Teelh 0.0 3.0 4.0 0.0 3.0 Pyloric Caeca 55.0 48.0 38.0 0.0 39.0 Ave II :::: 8.3 Ave I, = Character R B U6 16 U7 Scales Above Lat. Line 27.0 42.7 37.5 33 .1 41.5 Laleral Line Scales 130.0 144.2 175.0 0.0 155.5 Gill Rakers 20.5 18.5 20.0 x 18.0 Basibrallch ial Teeth 0.0 3.0 6.0 0.0 2.0 Pyloric Caeca 55.0 48.0 53.0 ( ill!: 40.0 Ave ]6= 26.1 Ave 17 = I, = 100 ' [(U,-B)i(R-B)] where, R :::: the mean of the character range for rainbow trout B = the adjusted mean of the character range for Bonneville cutlhroatlrout Ua :::: the raw character count for each fish I, = the character hybrid index for each fish Ave la :::: the average of the character hybrid indices for each fish 0 = Bonneville 30 - 70 ~ Ilybrid !"'o.Q .=j{lIil'b9W• _ L..JI I, 61.8 0.0 x 0.0 0.0 15.4 17 7.6 0.0 x 33.3 0.0 10.2 26 U3 I, U4 14 U, I, 36.0 , 42.7 37.0 36.3 35.5 ,:\5.9 157.5 0.0 166.0 0.0 160.5 0.0 19.0 x 18.0 x 18.0 x 4.0 0.0 2.0 33.3 1.0 66.7 43.0 0.0 40.0 0.0 44.0 0.0 Ave 13 = 10.7 Ave I, = 17.4 Ave 1,= 28.1 Us Is U, I, UIO 110 37.0 36.3 33.5 58.6 36.0 42.7 161.5 0.0 150.5 0.0 166.0 0.0 20.0 x 16.0 x 17.0 x 6.0 0.0 0.0 DM I, - U., 14 U, - I, - 23.6 38.0 29.9 42.0 4.5 40.5 14.0 0.0 169.5 0.0 170.5 0.0 172.5 0.0 x 18.0 x 17.0 x 18.0 x 0.0 10.0 0.0 1.0 66.7 15.0 0.0 0.0 42.0 0.0 55.0 r 100.0 46.0 0.0 5.9 Ave I, = 7.5 Ave I, = 42.8 Ave I, = 3.5 I, U, 1, U, I, U IO 1'0 23.6 40.5 14.0 4 1.5 7.6 36.5 39.5 0.0 183.0 0.0 175.5 0.0 163.5 0.0 x 18.0 x 18.0 x 17.0 x 0.0 4.0 0.0 5.0 0.0 3.0 0.0 0.0 45.0 0.0 53.0 71.'1 48.0 0 .0 5.9 Ave I, = 3.5 Ave I , = 19.8 Ave 110 = 9.9 Appendix B. Putative Bonneville cUl1hroat trout Population: Pctcctnect Cr.- RF Water 10: None (Spanish Fork Drainage) Population Average = 16.7 Character R B U, I, U, - . Scal es Above Lat. Line 27.0 42.7 41.0 10.8 36.0 Lateral Line Scales 130.0 144.2 172.0 0.0 154.5 Gill Rakers 20.5 18.5 19.0 x 2 1.0 Basibranchial Teeth 0.0 3.0 0.0 r 100.0 2.0 Pyloric Caeca 55.0 48.0 40.0 0.0 x Ave 11 = 27.7 Ave 12 == Character R B U6 16 U, Scal es Above La!. Line 27.0 42.7 41.0 10.8 43.0 Lateral Line Scales 130.0 144.2 169.5 0.0 200.0 Gill Rakers 20.5 18.5 20.0 x 18.0 Basibranchial Teeth 0.0 3.0 1.0 66. 7 2.0 Pyloric Caeca 55.0 48.0 x x 48.0 Ave 16 = 25.8 Ave 17 = I, ~ 100 *[(U"·B)/(R-B)] where, R = the mean of the character range for rainbow trout B ~ the adju sted mean of the character range for Bonnevill e cutthroa1 troLit Ua = the raw character COllnt for each fi sh la = the character hybrid index for each fi sh Ave Ia = th e average of the character hybrid indices for each fish 0 ::: Bonnev ille 30 · 70 = 11ybrid 110.0 :: l~i! \bOI\' _._-:_:::j I, . U, . ' I, . U, - I, - U, .. I, . 42.7 42.0 4.5 41.0 10.8 39.0 23 .6 0.0 188.0 0.0 185.0 0.0 177.5 0.0 x 19.0 x 18.0 x 19.0 x 33 .3 6.0 0.0 1.0 66.7 4.0 0.0 x 43.0 0.0 46.0 0.0 41.0 0.0 25.3 Ave 13 = 1.1 Ave ]4 :::0 19.4 A,rcls= 5.9 I, U, I, U9 I, UIO 110 - --- 0.0 37.5 33 .1 32.5 65.0 40.5 14.0 0.0 193.0 0.0 152.5 0.0 196.0 0.0 x 19.0 x 22.0 x 18.0 x 33.3 0.0 L 19 0.0 6.0 0.0 4.0 0.0 0.0 37.0 0.0 41.0 0.0 38.0 0.0 8.3 Ave Is = 33.3 Ave 19 = 16.2 Ave 110 = 3.5 28 Appendix 13. Putative l30nncvillc cuUhroattrout Population: Wimmer Ranch Creel, Water 10: nOlle g l ven Population Average = 12.6 Character R B U, I, U, - Scales Above l.at. Lille 270 42.7 37.5 33. 1 42.5 Lnlcml Line SC(llcs 130.0 144 .2 192.0 0.0 I R 1.5 Gill Rakers 20.5 IS .5 20.0 x I c).0 Basibranchial 'J'eeth 0.0 3.0 3.0 0.0 4.0 Pyloric Caeca 55.0 '18 .0 42 .0 0.0 43.0 Ave II ::: 8.3 Ave 12 ::: C hrtrctCICr R B U" I, U, Sca les Above Lat. Lille 270 42.7 36.0 ·12.7 39.0 Lateral Line Scales 130.0 144.2 154.0 0.0 17S.0 Gil l Rakers 20.5 IS. 5 19.0 x I 'J.O l3as ibranchial Teeth 0.0 3.0 4 .0 0.0 8.0 Pyloric Caecn 55.0 48 .0 43.0 0.0 41.0 Ave 1,,= 10.7 Ave I, = I" "' 100 *1(U,,-ll)/(R-I3)] where. R ~'- the mean of th e chnracler range 1'01' rainbow trout 13 :: the adjusted mean of tile character range for l10nncvillc ClIlthronl trout U~ =-- the rnw character count for each fi sh I ~ == the ch.uC\ctcr hybrid index for each !Ish Ave I ~ = the average of the character hybrid illdices for each fish 0 =" J1onn('v il lc 30 - 70 ' II ) h, it! 100 1(";lIllow 29 I, U, 11 U, I, U, I, 0.0 36.0 ·12.7 35.0 '19.0 39.0 23.6 0.0 15 1.0 0.0 159.0 0 .0 165 .5 0.0 x 20.0 x IS .O x 17.0 x 0.0 2.0 33.3 4.0 0.0 0.0 100.0 0.0 x x 39.0 0.0 44.0 0.0 lUI Ave 13 =: 25.3 Ave 101 = 12.3 Ave I~ = ]0.'1 I, Us I, U, I, U,O 110 23. 6 40 .5 14 .0 39.5 20.4 39.0 23.6 0.0 170 .5 0.0 166.0 0.0 159.0 0.0 x 19.0 x 20.0 x 18.0 x 0.0 7.0 0.0 1.0 66. 7 x x 0.0 35.0 0.0 38.0 0.0 37.0 0.0 5.9 Ave Is = 3.5 Ave 1'1 = 21.8 Ave 110 == 7.9 Appendix B. Putative Bonneville cuttlu·oat trout Population: WaterlD: Population Average = Character Scales Above Lat. Line Lateral Line Scales Gill Rakers Basibranchial Teeth Pyloric Caeca Character Little Deer Creek - South Fork V AF 040A 01 14.9 R B v, I, 27.0 42.7 41.0 10.8 130.0 144.2 187.0 0.0 20.5 18.5 19.0 x 0.0 3.0 20.0 0.0 55.0 48 .0 40.0 0.0 Ave I, ~ 2.7 R B U, I, v, I, 37.5 33. 1 155.0 0.0 19.0 x 0.0 1:.1 0'1..0 36.0 0.0 Ave I, ~ 33.3 U, I, ._-- ---_ ... _- -- - _.- --- Scales Above Lat. Line 27.0 42.7 34 .0 55.4 35.0 Lateral Line Scales 130.0 144.2 162.0 0.0 174.5 Gill Rak ers 20.5 18.5 19.0 x 19.0 Basibranchial Teeth 0.0 3.0 2.0 33 .3 4.0 Pyloric Caeca 55.0 48.0 34.0 0.0 39.0 AvcI,~ 22.2 Ave I, ~ I, ~ 100 *[(U,-B)/(R-B)] where, R = the mean of the character range for rainbow trout B ~ the adjusted mean of the character range for Bonneville cutthroat trout Ua = the raw character count for each fish 13 = the character hybrid index for each fi sh Ave I, ~ the average of the character hybrid indices for cach fi sh 49.0 0.0 x 0.0 0.0 12.3 30 U) I, U4 I., Vs Is 36.5 , 39.5 36.5 39.5 38.5 26.8 172.5 0.0 146.0 0.0 162.0 0.0 20.0 x 20.0 x 19.0 x 5.0 0.0 3.0 0.0 2.0 33.3 40.0 0.0 42.0 0.0 40.0 0.0 Ave I, ~ 9.9 Ave 14 ~ 9.9 Ave Is~ 15.0 v, I , U. I. U,o 1'0 . - -- 38.5 26.8 40.5 14.0 37.5 33.1 166.0 0.0 169.5 0.0 170.5 0.0 20.0 x 20.0 x 18.0 x 0.0 rJQM~ 11 .0 0.0 6.0 0.0 44.0 0.0 37.0 0.0 43.0 0.0 Ave I, ~ 31.7 Ave I. ~ 3.5 Ave 110 = 8.3 Appendix B. Putative Bonneville cutthroat trout Population: Water ID: Little Diamond Creek V AK 020B 01 ]'opulatioll Average = 25.2 Character R B u, I, u, I, ---- -- --------- Sca les Above Lat. Line 27.0 42.7 32.5 65.0 39.0 Lateral Line Scales 130.0 144.2 178.5 0.0 155.5 Gi ll Rakers 20 .5 18.5 20.0 x 18.0 Basibranchial Teeth 0.0 3.0 3.0 0.0 4.0 Pyloric Caeca 55.0 48.0 47.0 0.0 36.0 Ave I] = 16.2 Ave '2 = Characler R B u. I. u, Sca les Above Lat. Line 27.0 42.7 41.5 7.6 37.5 Lateral Line Scales 130.0 144.2 164.5 0.0 165.0 Gill Rakers 20.5 18.5 19.0 x 19.0 Oasibranchial Teeth 0.0 3.0 3.0 0.0 0.0 Pyloric Caeca 55.0 48.0 39.0 0.0 31.0 Ave I. = 1.9 Ave I , = I, = 100 *[(U,-B)/(R-B)] where, R = the mean of the character range for rainbow trout B = Ihe adjusled mean of the characler range for Bonneville cUlIhroat troul Ua = the raw character count for each fi sh I, = Ihe characler hybrid index for each fi sh Ave I, = Ihe average of Ihe character hybrid indices for each fi sh 0 = Bonneville 30 - 70 = Ilybrid 100 Q Rainbow 23.6 0.0 x 0.0 0.0 5.9 I, 33.1 0.0 x [ IOO.Q 0.0 33.3 31 U3 13 U4 14 u, I, _. 35.0 49.0 32.5 65.0 35.5 45 .9 167.5 0.0 15 1.5 0.0 161.0 0.0 18.0 x 18.0 x 19.0 x 3.0 0.0 0.0 LlOo:O' 5.0 0.0 36.0 0.0 44 .0 0.0 37.0 0.0 Ave I, = 12.3 Ave 14 = 41.2 Avels = IJ.s u, I, u, I, u" 1'0 32.0 68.2 29.5 l 84.1 36.0 42.7 147.5 0.0 143 .0 8.5 170.5 0.0 20.0 x 17.0 x 19.0 x 0.0 tToo.O 0.0 [[.I) "0 4.0 0.0 55.0 100.0 49.0 14.3 36.0 0.0 Ave I, = 67.0 Ave I, = 51.7 Ave 1'0 = 10.7 Appendi x B. Putati ve Bonneville cutthroat trout Populati on: Water I]): POJlulation Average = Character Sca les Above Lal. Line Lateral Line Scales Gill Rakers Basibranch ia l Tecth Pyloric Caeca Character Scales Above Lat. Line Lateral L ine Sca les Gill Rakers Basibranchia l "J'ecth Pyloric Caeca I, = 100 *[(U,-B)/(R-I3)] Cottonwood Creek V AK 020G 01 2J.J R 13 U, 270 42.7 37.0 130.0 144.2 157 .0 20.5 18.5 19.0 0.0 3.0 0.0 55.0 48.0 46.0 Ave I, = R B U, 27.0 42 .7 35.5 130.0 144.2 157 .0 20.5 18.5 19.0 0.0 3.0 3.0 55.0 48.0 35.0 Ave 16= w here, R = the mean of the character range for rainbow trout I, U, 36.3 37 .5 0.0 156.5 , 18.0 f lQOO 2.0 0.0 43.0 3~ .1 Ave I, = I, U, 45.9 37.5 0.0 141 .0 x 18.0 0.0 0.0 0.0 35.0 11.5 Ave 17 = B = the adjusted mean of the character range for Bonneville cutthroat trollt u~ == the raw character count for each fi £h I, = the character hybrid index for each lish A ve III = the average of the character hybrid indices for each fish 0 = Bonn evill e 30 - 70 = Ilybrid ~.I 00 ~".!!.lbow I, U3 I, U" I" U, I, 33 .1 36.0 42.7 37.5 33. 1 38.0 29.9 0.0 151.0 0.0 141.0 22.5 160 .5 0.0 , 18.0 x 19.0 x 17.0 x 33.3 0.0 1 100.0 7.0 0.0 4.0 0.0 0.0 36.0 0.0 36.0 0.0 28.0 0.0 16.6 Ave I, = 35.7 Ave I" = 13.9 Ave 15 = 7.5 I, U, I, U9 19 U IO 110 33.1 3 1.5 I 71.3 38.0 29.9 30.5 I J2] 22.5 165.0 0.0 144.5 0 .0 150.5 0.0 x 18.0 x 19.0 x 18.0 x 100,9 2.0 33.3 3.0 0.0 4.0 0.0 0.0 33 .0 0.0 4 1.0 0.0 37.0 0.0 38.9 Ave Is = 26.2 Ave 19 = 7.5 Avc11O = 19.4 32 Append ix B. Putalive Bonnevill e cutthroat trout Population: Water 10: Population Average = Character Scates Above Lat. Line Lateral Line Scales Gi ll Rakers Basibranchial Teeth Pylo ric Caeca Character Scales Above LaC Line Lateral Line Scales Gill Rakers Basibranch ial Teeth Py loric Caeca I, ~ 100 '[(U,-B)I(R-il)] Clear Creek - RF V AK 04011 01 8.5 R B 27.0 42.7 130.0 144.2 20 .5 18.5 0.0 3.0 55.0 48.0 R B 27.0 42.7 130.0 144.2 20.5 18.5 0.0 3.0 55.0 48.0 U, 40.0 176.0 20.0 3.0 39.0 Ave II = U, 38.5 178 .5 18.0 9.0 37.0 Ave 16= where, R ::.::. the mean of the character range for rainbow trout I, U, 17.2 47.5 0.0 194.5 x 2 1.0 0.0 5.0 0.0 38.0 4.3 Ave 12 ::: 16 U, 26.8 42.0 0.0 177.0 x 19.0 0.0 7.0 0.0 35.0 6.7 Ave I, ~ B = the adj usted mea n of the character range for Bonnevill e cutthroat trout Ua = the raw character count for each fi sh la = the character hybrid index for each fi sh Ave In = the average of tile character hybrid indices for each fi sh 0 = !lonneville 30 - 70 = Ilybrid I J 00 I(ainhuw 33 I, U, I, U, I, Us I, 0.0 40.0 17.2 37.5 33. I 34.5 52.2 0.0 184.0 0.0 165.5 0.0 175.5 0.0 x 19.0 x 19.0 x 18.0 x 0.0 I 1.0 0.0 3.0 0.0 2.0 33.3 ! 0.0 4 1.0 0.0 43.0 0.0 38.0 0.0 , 0.0 Ave IJ = 4.3 Ave 14 = 8.3 Ave Is= 21.4 I, U, I, U. 19 U IO 110 4.5 35.5 45.9 37.5 33 . I 36.0 42.7 0.0 170.5 0.0 178.5 0.0 174.5 0.0 x 20.0 x 17.0 x 19.0 x 0.0 6.0 0.0 2.0 33.3 5.0 0.0 0.0 48.0 0.0 46.0 0.0 37.0 0.0 1.1 Ave I, ~ 11.5 Ave I, ~ 16.6 Ave ILo = 10.7 Appendix B. Putative Bonneville cutthroat trout Populati on : Water 10: Deep Creel{ VI AA 5l0G Population Average = 14.8 Character R B Sca les Above Lat. Line 27.0 42.7 Lateral Line Scales 130.0 144.2 Gi ll Rakers 20 .5 18.5 Basibranchial Teeth 0.0 3.0 Pyloric Caeca 55.0 48.0 Character R B Scales Above Lat. Line 27.0 42.7 Lateral Line Sca les 130.0 144.2 Gi ll Rakers 20.5 18.5 Basibranchial Teeth 0.0 3.0 Pyloric Caeca 55.0 48.0 I, = 100 *[(U,-B)/(R-Bl] U, 41.5 163.5 20.0 6.0 42.0 Ave I, = U, 35.5 147.0 19.0 4.0 41.0 Ave 16 = where, R = the mean of the character range for rainbow trout I, U2 7.6 38.5 0.0 161.5 x 20.0 0 .0 3.0 0.0 41.0 1.9 Avel, = 16 U7 45.9 38.0 0.0 161.0 x 20.0 0.0 2.0 0.0 41.0 11.5 Ave 17 = B = the adjusted mean of the character range for Bonneville cUllhroat trout Ua = the raw character count for each fish I, = thc character hybrid index for each fish A ve I, = the average of the character hyhid indices for each fish 0 = Bonneville 30 - 70 = Ilybrid Ii 00 ~ Rainbow ...... ""_ .... I, U3 13 U4 I., U, I, 26.8 38.5 26.8 36.0 42.7 38.5 26.8 0.0 156.0 0.0 161.0 0.0 159.0 0.0 x 20.0 x 19.0 x 19.0 x 0.0 6.0 0.0 0.0 ~ .LOO.O~ 4.0 0.0 0.0 41.0 0.0 37.0 0.0 42.0 0.0 6.7 Ave I, = 6.7 Ave 14 = 35.7 Ave Is= 6.7 17 U, I, U, I, U IO 110 29.9 38.5 26.8 34.0 55.4 37.0 36.3 0.0 156.5 0.0 151.0 0.0 149.5 0.0 x 2 1.0 x 20.0 x 20.0 x 33 .3 2.0 33.3 0.0 U..QQ.JL 5.0 0.0 0.0 42.0 0.0 37.0 0.0 41.0 0.0 15.8 Ave I, = 15.0 Ave I, = 38.9 AveI 1o = 9.1 34 Appendix B. Putative Bonneville cutthroat trout Population: Water 10: Population Average = Character ------ -- - - Scales Above Lat. Line Lateral Line Scales Gill Rakers Basibranchial Teeth Pyloric Caeca Charact.er Scales Above Lat. Line Lateral Line Scales Gill Rakers Basibranchial Teeth Py loric Caeca I, = 100 *[(U,-B)/(R-D)] Birch Creek VI AB 050A 2 15.6 R B 27.0 42.7 130.0 144.2 20.5 18.S 0.0 3.0 55.0 48.0 R B 27.0 42,7 130 ,0 144 .2 20.5 18.5 0.0 3,0 55.0 48,0 U, 33.0 152.5 20.0 7,0 47.0 Ave I, = U, 34.5 151.5 19~0 8.0 42,0 Ave I, = where, R = the mean of the character range for rainbow trout I, U, 61.8 33.S 0.0 153.2 x 18~0 0.0 11.0 0.0 35,0 15.4 Ave 12 = 16 U, 52.2 31.0 0.0 151.0 x 18.0 0,0 11 .0 0,0 39.0 13.1 Ave I, = B = the adj usted mean of the character range for Bonneville cutLhroat trout Ua = the raw character count for each fish I, = the character hybrid index for each lish Ave 1ft = the average of the character hybrid indices for each fi sh 0 = Bonneville 30 - 70 lJybrid 100 ' I~ainbow . • I, U, I, U, I, U, I, ~--.------- 58.6 34 .S 52.2 35.0 49.0 35.S I 45 .9 0.0 150.S 0.0 151.0 0.0 15 1.0 0.0 x 18.0 x 20.0 x 20.0 x 0.0 12.0 0.0 16.0 0.0 4.0 O~O 0.0 41.0 0.0 43.0 0.0 42.0 0.0 14.6 Ave I, = 13.1 Ave 14 = 12.3 Avei,= 11.5 I, U, I, U, I, UIO 110 I 74.5 31.0 L 74.5 33.0 61.8 33.0 61.8 0.0 155.0 0.0 149.0 0.0 166.5 O~O x 18.0 x 2 1.0 x 20.0 x 0.0 2.0 33.3 5.0 0.0 8.0 0.0 0,0 43 .0 0,0 37.0 0.0 38.0 0.0 18.6 Ave I, = 27.0 Ave 19 = J5.4 Ave 110 = 15.4 35 Appendix B. Putative Bonneville cutthroat (rout Population: North Creel, Lower North Fork Water ID: VI AB 070 A Population Average = SUI Character R B u, I, u, I, - - Scales Above Lat. Line 27.0 42.7 30.5 l 7Ll 37.5 33. 1 Lateral Line Scales 130.0 144.2 144.0 1.4 148.5 0.0 Gill Rakers 20.5 18.5 18.0 x 16.0 x Basibranchial Teeth 0.0 3.0 • x 0.0 [ 100-:0 Pyloric Caeca 55.0 48.0 34.0 0.0 57.0 IQO.O Ave I, = 26.4 Ave I, = 58.3 Character R B U, I, U7 17 Scales Above Lat. Line 27.0 42.7 39.5 20.4 30 .0 1 80.9 Lateral Line Scales 130.0 144.2 179.5 0.0 138.0 43 .7 Gill Rakers 20.5 18.5 17.0 x 17.0 x Basibranchial Teeth 0 .0 3.0 0.0 CIQQJl_ 0.0 [I 00 .0~ Pyloric Caeca 55.0 48.0 38.0 0.0 38.0 0.0 Ave 1, = 30.1 Ave 17 = 56.1 I, = 100 *[(U.-B)/(R-B)] where, R = the mean of the character range for rainbow trout B = the adjusted mean of the character range for Bonneville cutthroat trout Ua = the raw character count for each fi sh la = the character hybrid index for each fi sh Ave I. = the average of the character hybrid indices for each fish 0 = Bonneville 30 - 70 = Ilybrid LQQ~b_olV ____ ~ 36 U3 13 u, I, u, I, - 28.0 L 93 .6 29.0 , 87.3 27.5 U6.8 ~ 138.5 40.1 136.5 . 54.2 148.0 0.0 18.0 x 18.0 x 19.0 x 0.0 [;&0.0 0.0 100.0 0.0 ~O 44.0 0 .0 53.0 71.4 57.0 100.0 Ave I, = 58.4 Ave I, = 78:2 Ave 1,= 74.2 Us Is U9 19 U IO 110 31.5 I 71.3 28.5 I 90A 32.0 . § 8.2 136.5 . 54.2 140.0 29.6 140.0 29.6 18.0 x 17.0 x 19.0 x 5.0 0.0 0.0 OQ9.&. 0.0 D:QQ;9 .. 33.0 0.0 34.0 0.0 38.0 0.0 Ave Is = 31.4 Ave 19 = 55.0 Ave 110 = ~9.4 Appendix B . Putative Bonneville cutthroat trout Population: Water ID: NF-North Creek-Middle Station VI AB 070A ])opulation Average = 16.7 Character . - .. - --_. -- R B U Scales Above Lat. Line 27.0 42.7 38.5 26.8 Lateral Line Scales 130.0 144.2 157.0 0.0 Gill Rakers 20.5 18.5 20.0 x Basibranchial Teeth 0.0 3.0 * x Pyloric Cacca 55.0 48.0 46.0 0.0 Ave I, ~ 8.9 Character R B U, I, Scal es Above Lat. Line 27.0 42 .7 33.5 58.6 Lateral Line Scales 130.0 144.2 152.5 0.0 Gill Rakers 20 .5 IS.5 19.0 x Basibranchial Teeth 0.0 3.0 0.0 ~ 100.0 Pyloric Caeca , 55.0 48.0 42.0 0.0 Ave I, ~ 39,6 I" ~ 100 *[CU"· B)/CR-B)] where, R = the mean of the character range for rainbow trout U2 - 36,S 145.0 18.0 0.0 40.0 Ave I, ~ U J 32.0 143.5 18.0 2.0 42.0 Ave 17 :::: B :::: the adjusted mean of the character range for I30nncville c utthroat trout Ua :::: the I'm\' character count for each fi sh I" ~ the character hybrid index for each fish A ve III :::: the average of the character hybrid indices for each fi sh o ~ Bonn eville 30· 70 ~ Hybrid 1100 ·' Rninbow ' < 39.5 0.0 x t L2M 0.0 34,9 I, 68.2 4.9 x 33.3 0.0 26.6 37 U U U , ' .' ., .. , , , 42.0 4 .5 41.5 7.6 41.5 7 .6 178.0 0.0 183.0 0.0 178.5 0.0 17.0 x 18.0 x 18.0 x 4.0 0.0 6.0 0.0 8.0 0.0 45.0 0.0 39 .0 0.0 43.0 0.0 Ave I, ~ 1.1 Ave I, ~ 1.9 Ave I, ~ 1.9 U, I, U9 I, U IO 1'0 39.0 23.6 40.5 14.0 37 .0 36.3 IS9.0 0.0 IS7.0 0.0 170.0 0.0 18.0 x 17.0 x 19.0 x 0.0 1 100.0 2.0 33.3 7.0 0 .0 46.0 0.0 44.0 0.0 42.0 0 .0 Ave I, ~ 30,9 Ave 19 = 11.8 Ave 110 :::: 9.1 Appendix B. Putative Bonneville cuttlu·oat trout Population: Water 10: North Fork-North Creek-Upper Station Vl AB 070 A Population Average = 9.7 Character R B U, I, U2 Scales Above Lat. Line 27.0 42.7 40.0 17.2 35.5 Lateral Line Scales 130.0 144.2 163.0 0.0 161.0 G ill Rakers 20.5 18.5 20.0 x 18.0 l3asibranchial Teeth 0.0 3.0 0.0 [ 100.0 5.0 Pyloric Caeca 55.0 48.0 43.0 0.0 36.0 Ave II = 29.3 Avc I, ~ Character R B U, 16 U, Sca les Above Lat. Line 27.0 42.7 40.5 14.0 41.0 Lateral Line Sca les 130.0 144.2 174.5 0.0 165.0 Gi ll Rakers 20.5 18.5 20.0 x * Basibranchial Teeth 0.0 3.0 6.0 0.0 11.0 Pyloric Caeca 55.0 48 .0 48.0 0.0 42.0 Avel,~ 3.5 Ave I, ~ I, ~ 100 *[(U,-B)/(R-B)] where, R :::: the mean of the character range for rainbow trout B ~ the adjusted mean of the character range for Bonneville cutthroat troul U3 = the Taw character count for each fi sh 13 = the character hybrid index for each fish Ave la = the average of the character hybrid indices for each fish o ~ Bonneville 30 - 70 = Hybrid I 00 ~ Rainbow 12 U, 13 U, 14 U, I, 45.9 40.0 17.2 38.5 26.8 38.0 29.9 0.0 165.5 0.0 171.5 0 .0 177.5 0.0 x 18.0 x 17.0 x 19.0 x 0.0 13.0 0.0 10.0 0.0 10 .0 0.0 0.0 40.0 0.0 42.0 0.0 41.0 0.0 11.5 Ave I, ~ 4.3 Ave I, ~ 6.7 Ave Is ~ 7.5 I, U, I, U, I, U IO 110 10.8 39.0 23.6 39.0 23.6 40.5 14.0 0.0 165.5 0.0 167.0 0.0 171.5 0.0 x 18.0 x 19.0 x 18.0 x 0.0 9.0 0.0 1.0 66.7 3.0 0.0 0.0 40.0 0.0 43.0 0.0 4 1.0 0.0 2.7 Ave I, ~ 5.9 Ave I, ~ 22.6 Ave 110 ~ 3.5 38 Appendix C 39 Appendix C. Putati ve Colorado River cutthroat trout Population: Water ID: East Fork - Boulder C"cek I AJ 11 OC Population AVCJ"agc = 1.2 Character R C u, Scales Above Lat. Line 27.0 42.7 45.5 Latera l Lin e Sca les 130,0 187.0 188.0 Gi ll Rakers 20,S 19.0 19.0 Basibranehial Teeth 0.0 3,0 6.0 Pyloric Caeca 55.0 41.0 41.0 Ave I, = Characte r R C U, Scales Above Lat. Line 27.0 42. 7 52.0 Latera l Line Sca les 130,0 187.0 199,5 Gi ll Rakers 20.5 19.0 19,0 Basibranchia l Teeth 0.0 3.0 10,0 Pylor ic Caeca 55.0 4 1.0 40.0 Ave I, = I, = 100 * [(U,·C)I(R·C)] where, R = the mcan o f the character range for ra inbow trout I, u, 0.0 46.5 0.0 184.5 x 18.0 0.0 5,0 0.0 38.0 0.0 Ave 12 = 16 U, 0.0 47.5 0.0 197 .5 x 19.0 0.0 6.0 0.0 * 0.0 Ave I, = C = the adjusted mean of the character range for Colorado Ri ver cuUhroat trout U3. = the raw character count for each fi sh la = the character hybrid index for each fi sh Ave la = th e average of the character hybrid indices for each fi sh 0 = Co lorado River 30 . 70 = Ilybrid J 00 .- Rainbow I, u, I, U, I, U, I, 0.0 49.0 0.0 44.0 0.0 48.0 0.0 4.4 206.0 0.0 192.5 0,0 194.5 0.0 x 19 ,0 x 18.0 x 18,0 x 0.0 7,0 0.0 10 ,0 0.0 4 .0 0.0 0,0 43 .0 14 .3 43.0 14.3 40.0 0.0 1.1 Ave I, = 3.6 Ave 14 = 3.6 Avc ls = 0.0 I, U, I, U9 19 U IO 110 0.0 47.5 0.0 45.5 0.0 42.5 1.3 0.0 183.5 6. 1 183.0 7,0 191.5 0.0 x 19.0 x 18.0 x 20.0 x 0.0 9,0 0.0 14.0 0.0 6,0 0.0 x 33,0 0.0 41 ,0 0.0 41.0 0 .0 0.0 Ave I, = 1.5 Ave 111 = 1.8 Avc11O = 0.3 40 Appendix C. Putative Colorado River cutthroat trout Population: Water ID: Indian C"cel< I BG 03 Population Avcrage = 22.4 Charac ter R C U, I, U, I, - --- ---- ----_ .. Scales Above Lat. Line 27.0 42 .7 38.0 29.9 38.5 Lateral Line Scales 130.0 187.0 170.5 28 .9 17 1.0 Gi ll Rakers 20.5 19.0 18.0 x 20.0 Basi branchial Teeth 0.0 3.0 4.0 0.0 0.0 Pyloric Caeca 55.0 4 1.0 4 1.0 0.0 37.0 Ave I, ~ 14.7 Ave I, ~ Character R C U6 I, U7 Scales Above Lat. Line 27.0 42.7 40.0 17.2 39.0 Lateral Li ne Scales 130.0 187.0 169.5 30.7 154.0 Gill Rakers 20.5 19.0 19.0 x 2 1.0 Basibranchial Teeth 0.0 3.0 9.0 0.0 9.0 Pyloric Caeca 55.0 41.0 42.0 7.1 35 .0 Ave I, ~ 13.8 Ave 17 ~ I, ~ 100 *[(U,-C)/(R-C)] where, R = the mean of the character nmge for rainbow trout C ~ the adjusted mean of the character range for Colorado River cutthroat trout Ua = the raw character count for each fi sh I, ~ the character hybrid index for each !ish Ave I, ~ the average of the character hybrid indices for each !ish 0 = Colorado River 30 - 70 ~ Ilybrid 100 Rainbow 26.8 28. 1 x i IQQ)~ 0.0 38.7 17 23.6 57.9 x 0.0 0.0 20.4 41 U, I, U, I., U, I , - -- - ------- - - - - --- 35.5 45.9 37.0 36.3 35.0 49.0 171.5 27.2 175.5 20.2 168.0 33.3 20.0 x 19.0 x 21.0 x 6.0 0.0 3.0 0.0 0.0 l\Oo-:O~ 29.0 0.0 38.0 0.0 36.0 0.0 Ave I, ~ 18.3 Ave I, ~ 14.1
geography
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batch_date
Tue, 05 May 2020 19:04:31 +0000
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https://digitallibrary.utah.gov/awweb/awarchive?type=download&item=66149
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Content: 59e7e4ac1e474f70dc7eb283d560cc2bd5f75a4f | Abstract: a34d1afe417c4f3777e78a567e86d5e40df372f6